My best guess according to this resource: https://www.researchgate.net/publication/365417294_Considerations_on_the_Systematics_of_Neotropical_Pierina_with_the_Description_of_Two_New_Species_of_Phulia_Herrich-Schaffer_from_the_Peruvian_Andes_Lepidoptera_Pieridae_Pierinae_Pierini#fullTextFileContent
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